The actual formula for the shannonweiner diversity index is. Thus, when conducting simple statistical analyses of a specific effect of disturbance on diversity, the. Learn how to calculate shannon wiener diversity index with clear definition, formula and example. Demonstration of how to calculate the shannonwiener diversity index in microsoft excel. Natural resource biometrics measures of diversity shannon. A diversity index also called phylogenetic or simpsons diversity index is a quantitative measure that reflects how many different types such as species there are in a dataset a community and that can simultaneously take into account the phylogenetic relations among the individuals distributed among those types, such as richness, divergence or evenness. Relative to other diversity indices, such as simpsons index, it is considered sensitive to the. First, enter the number of species, and then enter the name you wish to give the species, if available, and the given populations for each of the speciesin any given o. The shannon index has been a popular diversity index in the ecological literature, where it is also known as shannon s diversity index, the shannon wiener index, the shannon weaver index and the shannon entropy. Relative to other diversity indices, such as simpsons index, it is considered sensitive to the addition of rare species krebs, 1989.
In implan, the sw index for a region is calculated as follows. Note that lower values indicate more diversity while higher values indicate less diversity. In chemical graph theory, the wiener index also wiener number introduced by harry wiener, is a topological index of a molecule, defined as the sum of the lengths of the shortest paths between all pairs of vertices in the chemical graph representing the nonhydrogen atoms in the molecule. A value of one perfect diversity occurs when all industries are present in the region, with employment spread equally among them. The shannon index, sometimes referred to as the shannonwiener index or the shannonweaver index 1, is one of several diversity indices used to measure diversity in categorical data. Woody species diversity and important value indices in dense dry forests in abdoulaye wildlife reserve togo, west africa hodabalo pereki1, kperkouma wala1, thomas thielclemen2, michael p. Shannon index of diversity is sometimes referred to as the shannonwiener index and on other occasions referred to as the shannonweaver index. This comparison can be between two different habitats or a comparison of one habitat over time. The shannon index is a measurement used to compare diversity between habitat samples. Richness and evenness the heterogeneity of a landscape can be quantified using the shannon measures for richness and evenness.
Pdf shannonwiener diversity index has been used extensively in environmental studies to estimate the species richness and abundance of. Regarding your result, a shannon wiener diversity index of 2. Diversity of classes shannon, simpson, species statsdirect. A greater number of species and a more even distribution both increase diversity as measured by h. Also does anyone know if and how they can be done using primer 5. Woody species diversity and important value indices in dense.
Used to compare diversity, doesnt give a measure of dominance. H the shannon diversity index p i fraction of the entire population made up of species i s numbers of species encountered. The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field. The shannon wiener diversity index was calculated for the total zooplankton community found in lake superior in 1973, as well as for its components ie. We suggest an explanation for this dual use of terms and in so doing we offer a tribute to the late claude shannon who passed away on 24 february 2001.
Choose this diversity calculation from the alpha diversity tab. Simpson index must also be calculated for this will give you a general idea regarding dominance. Diversity index from wikipedia, the free encyclopedia redirected from diversity index a diversity index is a statistic which is intended to measure the differences among members of a set consisting of various types of objects. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. Application of shannons index to study diversity with. Aug 28, 2014 in analyses using only one diversity index, similar significant effects of lui would have been detected using any of the indices we included, except e. The two aims of this paper have something in common. Measurements of biodiversity marinespecies introduced. The shannonweiner diversity index is a common way of showing that diversity involves not only numbers of different species, but also how well each of these species is represented in different habitats. Shannons diversity index real statistics using excel. Pdf on jan 1, 2006, kathleen a nolan and others published beachcomber biology. Pdf the shannon diversity index has long been used extensively in marine ecological studies in coral reef ecosystems.
The actual formula for the shannon weiner diversity index is. Some analysts use 1 eh so that higher values indicate higher diversity. A diverse species assemblage also contributes to ecosystem diversity. It is simply the information entropy of the distribution, treating species as. Learn vocabulary, terms, and more with flashcards, games, and other study tools. Species and biological diversity choices of diversity. Better stories can be told about simpsons index than about shannons index, and still grander narratives about rarefaction hurlbert 1971.
A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot. Also, that shannonwiener diversity approaches log1d as p approaches 0. Relative to other diversity indices, such as simpsons index, it. Shannon index the idea behind this index is that the diversity of a community is similar to the amount of information in a code or message.
Instead we use a measure of the distribution of observations among the categories. If youd like more research, let me know and ill point you in the direction of the journals i used. This diversity measure is based on information theory. S, total number of species in the community richness. A diversity index of 65 translates to a probability of 65 percent that two people randomly chosen from the us population would belong to different race or ethnic groups. The importance of species diversity to the ecosystem sciencing. Shannon index is a commonly used diversity index that takes into account. Since calculating the wiener index of a graph can be computationally expensive, we give some cheaply computable lower and upper bounds for the wiener index given certain graph properties in chapter 3. Better stories can be told about simpsons index than about shannon s index, and still grander narratives about rarefaction hurlbert 1971. A diversity index is a mathematical measure of species diversity in a community. These values were then subjected to a model to develop statistically homogeneous zones, and to determine the temporal and spatial variability of the diversity values.
There are many methods that ecologists use to calculate species diversity. What test should i use with shannon diversity index values. Two commonly used measures simpsons index d s and shannon s index h. Piln pi where h is the index of diversity, pi is the importance value of a species as a proportion of all species. A diversity index is the measure of species diversity in a given community. The maximum diversity hmax of a sample is found when all species are equally abundant. A a diversity index is a mathematical measure of species diversity in a given community. The shannon index, sometimes referred to as the shannon wiener index or the shannon weaver index 1, is one of several diversity indices used to measure diversity in categorical data. Shannonwiener diversity index in excel microbial research.
Measuring diversity has been of historical significance and due to the obvious declines in habitat diversity this data is still valuable. A tribute to claude shannon 19162001 and a plea for. In our study we closely examine functional diversity indices to clarify their accuracy. It is simply the information entropy of the distribution, treating species as symbols and their relative population sizes as the probability. The most useful descriptions of diversity, therefore, present both measures of richness and evenness. Biodiversity calculator for the simpson and shannon indexes. C the more species you have, the more diverse the area. The sw index has been calculated and displayed by the implan data and software system for economic impact analysis since their 1999 data set. Species richness an index based on the number of species. If it is chosen as a diversity index, then all communities that share a particular value of shannon entropy are equivalent with respect to their diversity according to this index. Specifically, an index value of 1 means that all groups have the same frequency. What is the definition of shannonwiener diversity index.
Key wordsdiversity indices, shannon index, shannon wiener index, shannon weaver index, species richness, species diversity. It assumed that individuals are randomly sampled from an infinitely large community, and that all species are represented in the sample. These metrics are a measure of the likelihood that a selected pixel will have the same land cover class as the adjacent pixel. Its the likelihood that one species has an equal probability of being represented as all the other species.
Diversity indices that combine both richness and abundance i. In analyses using only one diversity index, similar significant effects of lui would have been detected using any of the indices we included, except e. We suggest that at the very least, authors should define. Hence, for a fixed simpson diversity index, d, the shannonwiener diversity index can take on at least all values greater than or equal to log1d. Jan 26, 2012 a diversity index is the measure of species diversity in a given community. Regarding your result, a shannonwiener diversity index of 2. Is shannonweiner species diversity index a reflection of. What sort of test should i use to get the pvalue after getting a shannon diversity index. How to calculate shannon wiener diversity index definition. Posts tagged shannon wiener index test difference between diversity. The shannonwiener diversity index is one measure that we will use to try. If i obtain the shannonweiner diversity index as 2.
If an ecosystem has poor species diversity, it may not function properly or efficiently. Thus, when conducting simple statistical analyses of a specific effect of disturbance on diversity, the choice of index does not appear particularly important. If it is chosen as a diversity index, then all communities that share a particular value of shannon entropy are equivalent with respect to. We will then consider one measure used to compare diversity among communities, jaccard.
Statistics shannon wiener diversity index tutorialspoint. According to the encyclopedia of earth, species diversity is a measurement of an ecosystems species richness and species evenness. This normalizes the shannon diversity index to a value between 0 and 1. This study calculates religious diversity index rdi scores for countries, regions and the world based on the shares of eight major world religions buddhists, christians, folk religions, hindus, jews, muslims, other religions considered as a group, and the religiously unaffiliated. Species richness and species diversity the terms diversity, species diversity and species richness are. If groups is given, finds the total number of species in each group see example on finding one kind of beta diversity with this option. The most widely used diversity index in the ecological literature is the shannon wiener diversity index. Here, we will explore two measures of biodiversity. Demonstration of how to calculate the shannon wiener diversity index in microsoft excel. Pdf modification of shannonwiener diversity index towards. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. B based on the species richness the number of species present and species abundance the number of individuals per species.
Jan 06, 2011 demonstration of how to calculate the shannon wiener diversity index in microsoft excel. Diversity indices provide more information about community composition than simply species richness i. Shannons diversity index for categorical data, there is no mean or median and so the measures of variation described in measures of variability cant be used. Diversity index from wikipedia, the free encyclopedia redirected from diversity index a diversity index is a statistic which is intended to measure the differences among members of a. As the overall behavior and consistency of functional diversity indices have never been described so far, the novice user risks choosing an inaccurate index or a set of redundant indices to represent functional diversity. Shannon wiener species diversity index formula data analysis. This calculator is free to use and is designed for biologists, ecologists, teachers, and students needing to quickly calculate the biodiversity indexes of an ecosystem. Those measures which are a cdiilbination of 5 and n total number of individuals, for instance the m2rgalefindex and the log series index a, also give these woods equal diversity table 4. The index is actually not a direct measure of species diversity. Statistics shannon wiener diversity index in the literature, the terms species richness and species diversity are sometimes used interchangeably. This presents the shannonwiener also incorrectly known as shannonweaver diversity index for. The shannonwiener diversity index was calculated for the total zooplankton community found in lake superior in 1973, as well as for its components ie. The addition of rare species to a sample causes only small changes in the value of d shannonwiener index also been called the shannon index and the shannonweaver index. The degree of uncertainty of predicting the species of a random sample is related to the diversity of a community.
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